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Characterization of habitats important for bryophytes
Mires, mountain streams and saxicolous habitats in temperate areas. Besides the components on bryophyte ecology and their patterns of distribution, this research has been focused also on the detection of bio indicator species as well as on the setting of predictive models to be used as tools for conservation purposes

Patterns of distribution of bryophyte species
Relating earth history events with evolution processes is one of my most recent research interests, particularly with Sphagnum. In this case I’ve been studying the molecular differentiation of Sphagnum populations as well as doing some preliminary work on the relations between populations of closely related Sphagnum taxa in Europe and Africa.
Phylogeography of Iberian deciduous oaks using molecular markers is another project I have been involved as principal investigator.

Conservation studies
Working mainly with the bryoflora of protected areas in Portugal. With this work many updates of the bryoflora were possible as well as the evaluation of the conservation status of protected taxa and populations. In addition, I contributed to the Iberian cartography and the Iberian Red List with the taxonomic revision and conservation of Sphagnum species in Portugal.

Rarity patterns and the causes for rarity
This is a ongoing research topic in collaboration with Prof. Lars Söderström. In this study, all taxa in a rather large liverwort family are classified along several rarity variables. An evaluation of their threat status (following the criteria of IUCN) is then compared with their rarity status, in addition to reproductive and dispersal parameters. In connection, I have been building up a taxonomical, nomenclatural and distribution database for all taxa of the genus Sphagnum that are recognized.

Most of the work is done in cooperation with Dr. Cecília Sérgio, Prof. Rui Figueira University of Lisbon, Prof. Rémy Petit (INRA, Bordeaux, France), and Prof. L. Söderström, NTNU, Norway, together with MSc and PhD students. Some of the results have been published in journals or presented in conferences